Bat viruses related to MERS-CoV

Prior to the emergence of MERS-CoV, a group of bat coronaviruses had been reported including Tylonycteris bat coronavirus HKU4 (BtCoV-HKU4) in Tylonycteris bats and Pipistrellus bat coronavirus HKU5 (BtCoV-HKU5) in Pipistrellus bats in China, E.isa/M/Spain/2007 in Eptesicus isabellinus bats in Spain and N.noc/VM366/2008/NLD in Pipistrellus pipistrellus bats in  the Netherlands.