In Africa, novel betacoronavirus related to SARS-CoV have been detected in Hipposideros and Chaerophon species from Ghana, Kenya and Nigeria. However, compared with Asian and European  SL-CoVs, these viruses of non-rhinolophid origin were phylogenetically distant to SARS-CoV. The Western African isolates even formed as a potential new lineage of Betacoronavirus in the phylogenetic tree.