SL-CoV were also discovered  in rhinolophids from Slovenia, Bulgaria and Italy in Europe These European SL-CoVs exhibited  significant genetic variation from China isolates. The strain BM48-31 from Rhinoluphus lasii in Bulgaria was highly  divergent from  Chinese isolates, displaying major sequence differences  in several genes including  ORF3b and ORF6 and lacking the coding region of ORF8 in its genome.