SL-CoV were also discovered in rhinolophids from Slovenia, Bulgaria and Italy in Europe These European SL-CoVs exhibited significant genetic variation from China isolates. The strain BM48-31 from Rhinoluphus lasii in Bulgaria was highly divergent from Chinese isolates, displaying major sequence differences in several genes including ORF3b and ORF6 and lacking the coding region of ORF8 in its genome.